Despite a range of ready-made nest boxes around Vane Farm at Loch Leven the bird has opted for the bin outside the visitors' centre door.
Luckily the nest was spotted by a visitor who saw the bird emerge from the box before it was used by a smoker.
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The RSPB said there were currently about four baby great tits in the nest.
Raymond Welsh, the visitor centre and coffee shop manager, said A couple on holiday from England told me that a tit was nesting in the cigarette box. "At first I thought they were mistaken and that the birds were just looking for food. Then I looked in the box and there she was sitting quietly on her nest.
"It's just great that nature is everywhere."
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